If the sub-populations are subject to different environmental variations (i.e. separated enough that a disaster could occur at one sub-population site without affecting the other sub-populations) but still allow individuals to travel between sub-populations, then the individual sub-populations are more likely to go extinct than the total population.
如果分居民都受到不同的环境变化（即分离不够，一个灾害可以发生在一个分人口的网站而不影响其他分人口），但仍然允许个人之间的旅行次人口，然后个人分居民更有可能灭绝于总人口。