At Manchester, Harden had studied the action of light on mixtures of carbon dioxide and chlorine, and when he entered the Institute he applied his methods to the investigation of biological phenomena such as the chemical action of bacteria and alcoholic fermentation.
在曼彻斯特，哈登研究了光在二氧化碳和氯混合时的作用，进入研究所后他用自己的方法来研究生物现象——比如细菌和酒精发酵的化学反应。