Working on cats, dogs, monkeys, and apes that had been bereaved of their cerebral hemispheres, he found that reflexes must be considered integrated activities of the total organism, not just the result of activities of the so-called reflex-arcs, a concept then generally accepted.
在研究被切除了半脑的猫、狗、猴及猩猩时，他发现反射必须考虑整个有机体的综合活动，不只是所谓的反射弧的活动结果，这一概念后来被普遍接受。